黄土丘陵区天然杜松种群结构及动态分析

doi:10.13866/j.azr.2023.08.11

干旱区研究 ›› 2023, Vol. 40 ›› Issue (8): 1304-1311.doi: 10.13866/j.azr.2023.08.11

黄土丘陵区天然杜松种群结构及动态分析

宋佳佳¹(),李钢铁¹(),郭靖捷¹,谷忠厚²,刘坤²,李治龙³,康霞⁴

1.内蒙古农业大学沙漠治理学院，内蒙古呼和浩特 010018
2.鄂尔多斯市林业和草原事业发展中心，内蒙古鄂尔多斯 017000
3.鄂尔多斯市林业和草原调查勘验中心，内蒙古鄂尔多斯 017000
4.伊金霍洛旗水利技术服务中心，内蒙古伊金霍洛旗 017200

收稿日期:2022-12-14 修回日期:2023-03-02 出版日期:2023-08-15 发布日期:2023-08-24
通讯作者: 李钢铁. E-mail: 13848817183@163.com
作者简介:宋佳佳（1997-），女，硕士研究生，主要从事沙区植物资源保护与利用. E-mail: hopesjj@163.com
基金资助:
鄂尔多斯市科技重大专项(2022EEDSKJZDZX012)

Population structure and dynamic analysis of natural Juniperus rigida in a loess hilly area

SONG Jiajia¹(),LI Gangtie¹(),GUO Jingjie¹,GU Zhonghou²,LIU Kun²,LI Zhilong³,KANG Xia⁴

1. Desert Science and Engineering College, Inner Mongolia Agricultural University, Hohhot 010018, Inner Mongolia, China
2. Ordos Forestry and Grassland Development Center, Ordos 017000, Inner Mongolia, China
3. Ordos Forestry and Grassland Survey and Inspection Center, Ordos 017000, Inner Mongolia, China
4. Ejin Horo Banner Water Conservancy Technology Service Center, Ejin Horo Banner 017200, Inner Mongolia, China

Received:2022-12-14 Revised:2023-03-02 Online:2023-08-15 Published:2023-08-24

摘要/Abstract

摘要：

以内蒙古黄土丘陵区天然杜松种群为研究对象，通过静态生命表、生存函数、动态量化分析以及时间序列模型分析天然杜松种群结构和动态变化。结果表明：幼苗个体数量居多，中树其次，成树较少，结合种群数量动态变化指数V_p_i>0，该种群则为增长型；存活曲线经曲线模型验证趋于Deevey-Ⅱ型，表明个龄级死亡率稳定；考虑外界干扰时动态指数趋于0，种群在干扰条件下增长不明显；生存函数分析种群表现为前期生存强中期稳定后期逐渐衰退的趋势；未来2~8各个龄级时间后，幼苗减少，中树和成树均增加；幼苗阶段对种群的更新和发展具有决定作用，建议对幼苗实施科学的保护措施，促进种群的更新。对杜松种群内在机制的研究，丰富了干旱区的植被建设，为杜松种群经营管理提供了参考，为黄土丘陵区植被的保护和恢复提供理论依据。

关键词: 黄土丘陵区, 杜松, 种群结构, 生存分析, 动态分析

Abstract:

This study focused on the natural Juniperus rigida population in the loess hilly region of Inner Mongolia. The structure and dynamic changes of the population were analyzed using a static life table, survival function, dynamic quantitative analysis, and time series model. The results showed that the population mainly comprised many individual seedlings, followed by medium and less mature trees. Combined with the dynamic change index V_pi > 0, the population was a growth type. The survival curve was verified using the curve model and tended to Deevey-II, indicating that the mortality rate of each age class was stable. Considering the external interference, the dynamic index tended to 0, and the population growth is not obvious under the interference condition. In the survival function analysis, the population showed a strong survival trend in the early stage, followed by a gradual decline in the middle stage. During the 2-to-8-year period, the number of seedlings decreased while middle-aged and adult trees increased. The seedling stage is crucial to the regeneration and development of the population. Therefore, implementing scientific protection measures for seedlings to promote population regeneration is recommended. Research on the internal mechanisms of the Juniperus rigida population can enrich vegetation construction in arid areas, provide a reference for population management, and provide a theoretical basis for vegetation protection and restoration in the loess hilly area.

Key words: loess hilly region, Juniperus rigida, population structure, survival analysis, dynamic analysis

宋佳佳, 李钢铁, 郭靖捷, 谷忠厚, 刘坤, 李治龙, 康霞. 黄土丘陵区天然杜松种群结构及动态分析[J]. 干旱区研究, 2023, 40(8): 1304-1311.

SONG Jiajia, LI Gangtie, GUO Jingjie, GU Zhonghou, LIU Kun, LI Zhilong, KANG Xia. Population structure and dynamic analysis of natural Juniperus rigida in a loess hilly area[J]. Arid Zone Research, 2023, 40(8): 1304-1311.

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参考文献 38

[1]	赵阳, 齐瑞, 焦健, 等. 尕海-则岔地区紫果云杉种群结构与动态特征[J]. 生态学报, 2018, 38(20): 7447-7457.
	[ Zhao Yang, Qi Rui, Jiao Jian, et al. The population structure and dynamic characteristics of Picea purpurea at the Gahai-zecha area[J]. Acta Ecologica Sinica, 2018, 38(20): 7447-7457. ]
[2]	卫波宁, 常子惠, 张永娟, 等. 雅鲁藏布江三大支流流域小花水柏枝种群结构及动态特征[J]. 生态学报, 2022, 42(24): 10241-10252.
	[ Wei Boning, Chang Zihui, Zhang Yongjuan, et al. Population structure and dynamic characteristics of Myricaria wardii in the basin of three branches of Yarlung Zangbo River[J]. Acta Ecologica Sinica, 2022, 42(24): 10241-10252. ]
[3]	杨彪, 张全建, 龚旭, 等. 雅砻江冬麻豆(Salweenia bouffordiana)种群结构与动态特征[J]. 生态学报, 2020, 40(4): 1184-1194.
	[ Yang Biao, Zhang Quanjian, Gong Xu, et al. Population structure and dynamic characteristics of Salweenia bouffordiana[J]. Acta Ecologica Sinica, 2020, 40(4): 1184-1194. ]
[4]	中国科学院中国植物志编委会. 中国植物志: 第7卷[M]. 北京: 科学出版社, 2004: 8-9.
	[ Editorial Committee of the Flora of China, Chinese Academy of Sciences. Flora of China:Vol.7[M]. Beijing: Science Press, 2004: 8-9. ]
[5]	王堃, 徐先存. 杜松精油成分分析及其抗焦虑作用研究[J]. 云南化工, 2022, 49(5): 58-60.
	[ Wang Kun, Xu Xiancun. Analysis of essential oil components and its anti-anxiety effect: A case study of Juniper essential oil[J]. Yunnan Chemical Industry, 2022, 49(5): 58-60. ]
[6]	王燕清. 杜松种子育苗技术[J]. 山西林业科技, 2022, 51(2): 46-47.
	[ Wang Yanqing. Seed seedling techniques of Juniperus rigida[J]. Shanxi Forestry Science and Technology, 2022, 51(2): 46-47. ]
[7]	张亚芳. 黄土高原不同地区杜松种群动态及繁殖特性研究[D]. 杨凌: 西北农林科技大学, 2014.
	[ Zhang Yafang. Study on Different Juniperus rigida Population Dynamics and Reproductive Characteristics of Loess Plateau[D]. Yangling: Northwest A & F University, 2014. ]
[8]	黄荣雁, 贯春雨. 杜松育苗及栽培技术[J]. 防护林科技, 2019, 37(11): 94-95.
	[ Huang Rongyan, Guan Chunyu. Cultivation technology of Juniperus rigida[J]. Shelter Forest Science and Technology, 2019, 37(11): 94-95. ]
[9]	李振勇. 冀北地区杜松的栽培与管理[J]. 河北林业科技, 2011, 39(2): 97.
	[ Li Zhenyong. Cultivation and management of Juniper in northern Hebei[J]. Hebei Forestry Publicity Center, 2011, 39(2): 97. ]
[10]	刘雨. 宁夏贺兰山杜松(Juniperus rigida)表型多样性与化感作用研究[D]. 杨凌: 西北农林科技大学, 2011.
	[ Liu Yu. Phenotypic Diversity and Allelopathic Effect of Juniperus rigida in Helan Mountain, Ningxia[D]. Yangling: Northwest A & F University, 2011. ]
[11]	苏日罕, 郭恩亮, 王永芳, 等. 1982—2020年内蒙古地区极端气候变化及其对植被的影响[J]. 生态学报, 2023, 43(1): 419-431. doi: 10.1016/j.chnaes.2022.07.008
	[ Su Rihan, Guo Enliang, Wang Yongfang, et al. Extreme climate change in the Inner Mongolia and their impacts on vegetation dynamics during 1982-2020[J]. Acta Ecologica Sinica, 2023, 43(1): 419-431. ] doi: 10.1016/j.chnaes.2022.07.008
[12]	刘建泉, 杨建红. 祁连圆柏种群结构和生活史分析[J]. 干旱区资源与环境, 2015, 29(7): 140-144.
	[ Liu Jianquan, Yang Jianhong. Life histories and structure of Sabina przewalskii population[J]. Arid Land Resources and Environment, 2015, 29(7): 140-144. ]
[13]	姜在民, 和子森, 宿昊, 等. 濒危植物羽叶丁香种群结构与动态特征[J]. 生态学报, 2018, 38(7): 2471-2480.
	[ Jiang Zaimin, He Zisen, Su Hao, et al. Population structure and dynamic characteristics of the endangered Syringa pinnatifolia Hemsl[J]. Acta Ecologica Sinica, 2018, 38(7): 2471-2480. ]
[14]	徐俏, 赵万羽, 魏岩, 等. 阿尔泰山东部林区森林种群结构和空间分布格局[J]. 干旱区研究, 2022, 39(6): 1885-1895.
	[ Xu Qiao, Zhao Wanyu, Wei Yan, et al. Forest population structure and spatial distribution pattern of different age tree species in forest area of eastern Altai Mountains[J]. Arid Zone Research, 2022, 39(6): 1885-1895. ]
[15]	卢杰, 郭其强, 郑维列, 等. 藏东南高山松种群结构及动态特征[J]. 林业科学, 2013, 49(8): 154-160.
	[ Lu Jie, Guo Qiqiang, Zheng Weilie, et al. Population structure and dynamic characteristics of Pinus densata in southeast Tibet[J]. Forestry Science, 2013, 49(8): 154-160. ]
[16]	董灵波, 马榕, 田栋元, 等. 大兴安岭天然林不同演替阶段共优势种种群结构与动态[J]. 应用生态学报, 2022, 33(8): 2077-2087. doi: 10.13287/j.1001-9332.202208.013
	[ Dong Lingbo, Ma Rong, Tian Dongyuan, et al. Structure and dynamics of co-dominant species succession stages of natural forests in Daxing’an Mountains, China[J]. Chinese Journal of Applied Ecology, 2022, 33(8): 2077-2087. ] doi: 10.13287/j.1001-9332.202208.013
[17]	吴承祯, 洪伟, 谢金寿, 等. 珍稀濒危植物长苞铁杉种群生命表分析[J]. 应用生态学报, 2000, 11(3): 333-336. pmid: 11767626
	[ Wu Chengzhen, Hong Wei, Xie Jinshou, et al. Life table analysis of Tsuga longibracteata population[J]. Chinese Journal of Applied Ecology, 2000, 11(3): 333-336. ] pmid: 11767626
[18]	李敏敏, 刘鹏程, 孔维民, 等. 濒危植物澜沧黄杉种群结构及动态特征[J]. 生态学报, 2022, 42(13): 5504-5515.
	[ Li Minmin, Liu Pengcheng, Kong Weimin, et al. Population structure and dynamic characteristics of the endangered Pseudotsuga forrestii Craib[J]. Acta Ecological Sinica, 2022, 42(13): 5504-5515. ]
[19]	王飞, 霍怀成, 赵阳, 等. 甘南高山林线岷江冷杉—杜鹃种群结构与动态[J]. 植物研究, 2019, 39(5): 664-672. doi: 10.7525/j.issn.1673-5102.2019.05.004
	[ Wang Fei, Huo Huaicheng, Zhao Yang, et al. Population structure and dynamics of original Abies faxoniana Rehd-Rhododendron simsii Planch in High-mountain timberline of southern Gansu Province[J]. Botanical Research, 2019, 39(5): 664-672. ] doi: 10.7525/j.issn.1673-5102.2019.05.004
[20]	陈晓德. 植物种群与群落结构动态量化分析方法研究[J]. 生态学报, 1998, 18(2): 104-107.
	[ Chen Xiaode. A study on the method of quantitative analysis for plant population and community structure dynamics[J]. Acta Ecologica Sinica, 1998, 18(2): 104-107. ]
[21]	肖宜安, 何平, 李晓红, 等. 濒危植物长柄双花木自然种群数量动态[J]. 植物生态学报, 2004, 42(2): 252-257. doi: 10.17521/cjpe.2017.0138
	[ Xiao Yi’an, He Ping, Li Xiaohong, et al. Sutdy on numeric dynamics of natural population of the endangered species Disanthus cercidifolius var. Longipes[J]. Chinese Journal of Plant Ecology, 2004, 42(2): 252-257. ] doi: 10.17521/cjpe.2017.0138
[22]	杨小平, 周龙, 蒋丽丽, 等. 野生樱桃李的种群统计与生存分析[J]. 干旱区资源与环境, 2020, 34(2): 154-160.
	[ Yang Xiaoping, Zhou Long, Jiang Lili, et al. Population statistics and survival analysis of Prunus divaricata[J]. Arid Land Resources and Environment, 2020, 34(2): 154-160. ]
[23]	张金峰, 葛树森, 梁金花, 等. 长白山阔叶红松林红松种群年龄结构与数量动态特征[J]. 植物生态学报, 2022, 46(6): 667-677. doi: 10.17521/cjpe.2021.0498
	[ Zhang Jinfeng, Ge Shusen, Liang Jinhua, et al. Population age structure and dynamics of Pinus koraiensis in a broadleaved Korean pine forest in Changbai Mountain, China[J]. Chinese Journal of Plant Ecology, 2022, 46(6): 667-677. ] doi: 10.17521/cjpe.2021.0498
[24]	李俊清, 祝宁. 红松的种群结构与动态过程[J]. 生态学杂志, 1990, 9(4): 8-12.
	[ Li Junqing, Zhu Ning. Population structure of Pinus koraiensis and its dynamics[J]. Chinese Journal of Ecology, 1990, 9(4): 8-12. ]
[25]	邱华, 舒皓, 吴兆飞, 等. 长白山阔叶红松林乔木幼苗组成及多度格局的影响因素[J]. 生态学报, 2020, 40(6): 2049-2056.
	[ Qiu Hua, Shu Hao, Wu Zhaofei, et al. Influencing factors of composition and abundance pattern of tree seedlings inbroad-leaved Korean pine (Pinus koraiensis) mixed forest, Changbai Mountain, China[J]. Acta Ecologica Sinica, 2020, 40(6): 2049-2056. ]
[26]	姚杰, 宋子龙, 张春雨, 等. 距离和密度制约对吉林蛟河阔叶红松林幼苗生长的影响[J]. 北京林业大学学报, 2019, 41(5): 108-117.
	[ Yao Jie, Song Zilong, Zhang Chunyu, et al. Effects of distance and density dependence on seedling growth in a broadleaved Korean pine forest in Jiaohe of Jilin Province, northeastern China[J]. Journal of Beijing Forestry University, 2019, 41(5): 108-117. ]
[27]	吴语嫣, 李守中, 孙眭涛, 等. 长汀水土流失区侵蚀劣地马尾松种群动态[J]. 生态学报, 2019, 39(6): 2082-2089.
	[ Wu Yuyan, Li Shouzhong, Sun Zitao, et al. Population dynamics of Pinus massoniana on eroded land in the soil erosion area of Changting County[J]. Acta Ecological Sinica, 2019, 39(6): 2082-2089. ]
[28]	刘鑫, 焦健, 王婷, 等. 巴丹吉林沙漠南缘沙地芦苇种群生态特征[J]. 干旱区研究, 2022, 39(1): 220-229.
	[ Liu Xin, Jiao Jian, Wang Ting, et al. Population ecological features of Phragmites australis in sandy habitats on the southern edge of Badain Jaran Desert[J]. Arid Zone Research, 2022, 39(1): 220-229. ]
[29]	王童犇, 朱芩, 侯晓巍, 等. 祁连圆柏群落特征沿年降水量梯度的变化格局[J]. 干旱区研究, 2021, 38(6): 1695-1703.
	[ Wang Tongben, Zhu Qin, Hou Xiaowei, et al. Community characteristics of Juniperus przewalskii along annual precipitation gradientst[J]. Arid Zone Research, 2021, 38(6): 1695-1703. ]
[30]	马丹丹, 库伟鹏, 夏国华, 等. 珍稀濒危植物堇叶紫金牛种群结构及动态分析[J]. 南京林业大学学报(自然科学版), 2021, 45(3): 159-164.
	[ Ma Dandan, Ku Weipeng, Xia Guohua, et al. Structure and dynamics of rare and endangered plant Ardisia violacea natural population[J]. Journal of Nanjing Forestry University(Natural Science Edition), 2021, 45(3): 159-164. ]
[31]	高洪治, 黄欣, 宿昊, 等. 秦岭两地区红桦种群结构与动态特征[J]. 北京林业大学学报, 2022, 44(9): 12-20.
	[ Gao Hongzhi, Huang Xin, Su Hao, et al. Structure and dynamic characteristics of Betula albo-sinensis populations in two regions in the Qinling Mountains of northwestern China[J]. Journal of Beijing Forestry University, 2022, 44(9): 12-20. ]
[32]	谭一波, 詹潮安, 肖泽鑫, 等. 广东南澳岛中华楠种群结构及动态特征[J]. 生态学杂志, 2010, 29(10): 1901-1906.
	[ Tan Yibo, Zhan Chao’an, Xiao Zexin, et al. Population structure and dynamic characteristics of Machilus chinensis in Nan’ao Island, Guangdong Province[J]. Chinese Journal of Ecology, 2010, 29(10): 1901-1906. ]
[33]	唐凤, 邹天才, 杨乃坤, 等. 稀有濒危植物贵州红山茶种群结构及数量动态变化的研究[J]. 广西植物, 2022, 42(3): 520-529.
	[ Tang Feng, Zou Tiancai, Yang Naikun, et al. Population structure and dynamics analysis of rare and endangered plant Camellia kweichowensis[J]. Guangxi Botany, 2022, 42(3): 520-529. ]
[34]	张锦堂, 潘志立, 田云海, 等. 云龙天池国家级保护区云南松种群年龄结构及动态分析[J]. 生态学报, 2022, 42(22): 9091-9099.
	[ Zhang Jintang, Pan Zhili, Tian Yunhai, et al. Age structure and dynamic of Pinus yunnanensis population in Yunlong Tianchi Nature Reserve[J]. Acta Ecologica Sinica, 2022, 42(22): 9091-9099. ]
[35]	杨艳波, 蔡起航, 巩合德. 滇中矮杨梅种群结构与动态特征[J]. 西部林业科学, 2022, 51(5): 146-152.
	[ Yang Yanbo, Cai Qihang, Gong Hede. Population structure and dynamic characteristics of Myrica nana in Central Yunnan[J]. Western Forestry Science, 2022, 51(5): 146-152. ]
[36]	张晓鹏, 于立忠, 杨晓燕, 等. 辽东山区天然更新红松幼苗种群结构与动态[J]. 应用生态学报, 2022, 33(2): 289-296. doi: 10.13287/j.1001-9332.202202.001
	[ Zhang Xiaopeng, Yu Lizhong, Yang Xiaoyan, et al. Population structure and dynamics of Pinus koraiensis seedlings regenerated from seeds in a montane region of eastern Liaoning Province, China[J]. Chinese Journal of Applied Ecology, 2022, 33(2): 289-296. ] doi: 10.13287/j.1001-9332.202202.001
[37]	谭菊荣, 袁位高, 李婷婷, 等. 极小种群野生植物细果秤锤树种群结构与动态特征[J]. 生态学报, 2022, 42(9): 3678-3687.
	[ Tan Jurong, Yuan Weigao, Li Tingting, et al. Population structure and dynamic characteristics of an extremely small population plant Sinojackia microcarpa[J]. Acta Ecologica Sinica, 2022, 42(9): 3678-3687. ]
[38]	彭闪江, 黄忠良, 彭少麟, 等. 植物天然更新过程中种子和幼苗死亡的影响因素[J]. 广西植物, 2004, 30(2): 113-121, 124.
	[ Peng Shanjiang, Huang Zhongliang, Peng Shaolin, et al. Factors influencing mortality of seed and seedling in plant nature regeneration process[J]. Guangxi Botany, 2004, 30(2): 113-121, 124. ]

x	A_x	a_x	l_x	lnl_x	d_x	q_x	L_x	T_x	e_x	K_x	S_x
Ⅰ	30	47	1 000	6.91	234.04	0.23	883	8058	3.16	0.27	0.77
Ⅱ	36	36	766	6.64	191.49	0.25	670	7175	2.97	0.29	0.75
Ⅲ	41	27	574	6.35	148.94	0.26	500	6504	2.80	0.30	0.74
Ⅳ	19	20	426	6.05	106.38	0.25	372	6004	2.60	0.29	0.75
Ⅴ	25	15	319	5.77	85.11	0.27	277	5632	2.30	0.31	0.73
Ⅵ	11	11	234	5.46	42.55	0.18	213	5355	1.96	0.20	0.82
Ⅶ	8	9	191	5.25	42.55	0.22	170	5143	1.28	0.25	0.78
Ⅷ	5	7	149	5.00	-	-	74	4972	0.50	-	-

黄土丘陵区天然杜松种群结构及动态分析

Population structure and dynamic analysis of natural Juniperus rigida in a loess hilly area

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径级	原始数据	M₂⁽¹⁾	M₃⁽¹⁾	M₄⁽¹⁾	M₅⁽¹⁾	M₆⁽¹⁾	M₇⁽¹⁾	M₈⁽¹⁾
Ⅰ	30
Ⅱ	36	33
Ⅲ	41	39	36
Ⅳ	19	30	32	32
Ⅴ	25	22	29	31	31
Ⅵ	11	18	19	24	27	27
Ⅶ	8	10	15	16	21	24	24
Ⅷ	5	7	8	13	14	19	21	22

[1]	肖旭, 郑诚, 丁成琴, 范陈哲, 白悦江, 林隆超, 闫婷, 高宇, 史海静. 基于ANUSPLIN的黄土丘陵区气象要素插值方法对比与评估[J]. 干旱区研究, 2023, 40(10): 1575-1582.
[2]	李浩,胡婵娟,赵荣钦,郭雷,满洲. 黄土丘陵区典型人工林的根系分布特征[J]. 干旱区研究, 2021, 38(5): 1420-1428.
[3]	董立国, 蒋齐, 蔡进军, 张源润, 许浩, 李生宝. 基于Biolog-ECO技术不同退耕年限苜蓿地土壤微生物功能多样性分析[J]. 干旱区研究, 2011, 28(4): 630-637.
[4]	赵维军, 许智超, 张岩, 朱岩, 朱清科. 半干旱黄土区沟间地浅沟分布特征[J]. 干旱区研究, 2011, 28(4): 586-591.
[5]	崔长美, 王孝安, 郭华, 李伟. 子午岭林区天然油松林的种群结构与动态[J]. 干旱区研究, 2011, 28(1): 111-117.

径级	原始数据	M₂⁽¹⁾	M₃⁽¹⁾	M₄⁽¹⁾	M₅⁽¹⁾	M₆⁽¹⁾	M₇⁽¹⁾	M₈⁽¹⁾
Ⅰ	30
Ⅱ	36	33
Ⅲ	41	39	36
Ⅳ	19	30	32	32
Ⅴ	25	22	29	31	31
Ⅵ	11	18	19	24	27	27
Ⅶ	8	10	15	16	21	24	24
Ⅷ	5	7	8	13	14	19	21	22

径级	原始数据	M₂⁽¹⁾	M₃⁽¹⁾	M₄⁽¹⁾	M₅⁽¹⁾	M₆⁽¹⁾	M₇⁽¹⁾	M₈⁽¹⁾
Ⅰ	30
Ⅱ	36	33
Ⅲ	41	39	36
Ⅳ	19	30	32	32
Ⅴ	25	22	29	31	31
Ⅵ	11	18	19	24	27	27
Ⅶ	8	10	15	16	21	24	24
Ⅷ	5	7	8	13	14	19	21	22